Southern Rockhopper References

Banks, J., A.V. Buren, Y. Cherel, and J. Whitfield.  2006. Genetic evidence for three species of rockhopper penguins, Eudyptes chrysocome. Polar Biology 30:61-67.

Bingham, M.  2002. The decline of Falkland Islands penguins in the presence of a commercial fishing industry.  Revista Chilena de Historia Natural 75:805–818.

BirdLife International 2009a.  Species factsheet: Eudyptes moseleyi. Downloaded from on 4 February 2009.

BirdLife International 2009b. Species factsheet: Eudyptes chrysocome. Downloaded from on 4 February 2009.

Boersma, P.D. and J. Parrish. 1999.  Limiting abuse: marine protected areas, a limited solution. Ecol. Econ. 31:287-304.

Boersma, P.D., D.L. Stokes and I.J. Strange. 2002.  Applying ecology to conservation:  tracking breeding penguins at New Island South Reserve, Falkland Islands. Aquatic Conservation Marine Freshwater Ecosystems 12:63-74.

Clausen, A. and N. Huin.  2003. Status and numerical trends of King, Gentoo, and Rockhopper penguins breeding in the Falkland Islands. Waterbirds 26(4):389-402.

Cooper, R.H.  1992. Rockhopper penguins at the Auckland Islands. Notornis 39:66-67.

Crawford, R.J.M., J. Cooper, B.M. Dyer, M.D. Greyling, N.T.W. Klages, D.C Nel, S.L. Peterson and A.C. Wolfaardt. 2003.  Decrease in numbers of the eastern Rockhopper penguin Eudyptes chrysocome filholi at Marion Island, 1994/95-2002/03. African Journal of Marine Science 25:487-498.

Crawford, R.J.M., J. Cooper, B.M. Dyer, M.D. Greyling, N.T.W. Klages, P.G. Ryan, S.L. Peterson, L.G. Underhill, L. Upfold, W. Wilkinson, M.S. De Villiers, S. Du Pleissis, M. Du Toit, T.M. Leshoro, A.B. Makhado, M.S. Mason, D. Merkle, D. Tshingana, V.L. Ward, and P.A. Whittington.  2003. Populations of surface-nesting seabirds at Marion Island, 1994/95-2002/03. African Journal of Marine Science 25:427-440.

Crawford, R.J.M., A.B. Makhado, L. Upfold, and B.M. Dyer.  2008. Mass on arrival of Rockhopper penguins at Marion Island correlated with breeding success. African Journal of Marine Science 30(1):185-188.

Cunningham, D.M. and P.J. Moors. 1994.  The decline of Rockhopper penguins (Eudyptes chrysocome)at Campbell Island, Southern Ocean and the influence of rising sea temperatures. Emu 94: 27-36.

Cuthbert, R.J., and E.S. Sommer.  2004. Population size and trends of four globally threatened seabirds at Gough Island, South Atlantic Ocean. Marine Orinthology 32:97-103.

Davis, L.S. and M. Renner. 2003.  Penguins. Yale University Press, United States.

Ellis, S.J.P., Croxall and J. Cooper. 1998. Penguin conservation assessment and management plan. IUCN/SSC conservation breeding specialist group, IUCN, Apple Valley.

Guinard, E., H. Weimerskirch, and P. Jouventin.  1998. Population changes and demography of the Northern Rockhopper penguin on Amsterdam and St Paul Islands. Colonial Waterbirds 21(2):222-228.

Hilton, G.M., D.R. Thompson, P.M. Sagar, R.J. Cuthbert, Y. Cherels, and S.J. Bury. 2006. A stable isotopic investigation into the cause of decline in a sub-Antarctic predator, the Rockhopper penguinEudyptes chrysocome. Global Change Biology 12:611-625.

Hindell, M.A. 1988. The diet of the rockhopper penguin Eudyptes chrysocome at Macquarie Island. Emu 88:227-233.

Hull, C.L., M. Hindell, K. Le Mar, P. Scofield, J. Wilson, and M.A. Lea. 2004.  The breeding biology and factors affecting reproductive success in Rockhopper penguins Eudyptes chrysocome at Macquarie Island. Polar Biology 27(11):711-720.

Huin, H. 2007. Falkland Islands Penguin Census 2005/2006. Downloaded from: on 12 February 2009.

IUCN 2008. 2008 IUCN Red List of Threatened Species. <>. Downloaded on 21 January 2009.

Jones, A.G., S.L. Chown, P.G. Ryan, N.J.M. Gremmen and K.J. Gaston. 2003. A review of conservation threats on Gough Island:  a case study for terrestrial conservation in the Southern Oceans. Biological Conservation 113:75-87.

Jouventin, P., R.J. Cuthbert, and R. Ottvall. 2006. Genetic isolation and divergence in sexual traits: evidence for the northern Rockhopper penguin Eudyptes moseleyi being a sibling species. Molecular Ecology 15:3413-3423.

Kirkwood, R., K. Lawton, C. Moreno, J. Valencia, R. Schlatter, and G. Robertson.  2007. Estimates of southern Rockhopper and Macaroni penguin numbers at the Ildefonso and Diego Ramirez Archipelagos, Chile, using quadrant and distance sampling techniques. Waterbirds 30(2):259-267.

Liljesthrom, M., S.D. Emslie, D. Frierson and A, Schiavini. 2008. Avian predation at a Southern Rockhopper Penguin colony on Staten Island, Argentina. Polar Biology 31:465-474.

Lynch, W. 1997. Penguins of the World. Firefly books, New York.

Marine Bio 2009. Rockhopper penguin. Downloaded from: on 29 January 2009.

Moors, P.J. 1986.  Decline in numbers of Rockhopper penguins at Campbell Island. Polar Record 23:69-73.

Muller-Schwarze, D. 1984. The Crested Penguins, pages 135-138 in The Behavior of Penguins. SUNY Press, Albany.

Murdoch, R. and P. Sagar. 2005. Rockhopper penguins (Eudyptes chrysocome filholi) foraging at Antipodes Islands. Notornis 52(2):75-80.

Oehler, D., W.R. Fry, L. Weakley Jr. and M. Marin.  2007. Rockhopper and Macaroni penguin colonies absent from Isla Recalada, Chile. The Wilson Journal of Orinthology 119(3):502-506.

Oehler, D., S. Pelikan, R. Fry, L. Weakley Jr., A. Kusch, and M. Marin. 2008.  Status of crested penguin (Eudyptes spp) populations on three islands in southern Chile. The Wilson Journal of Orinthology 120(3):575-581.

Penguin World 2009. Rockhopper Penguin. Downloaded from: on 18 February 2009.

Phelan, D. 1999. “Eudyptes chrysocome” (On-line), Animal Diversity Web. Accessed 9 February 2009 at

Poisbleau, M., L. Demongin, I. Strange, H. Otley, and P. Quillfeldt. 2008. Aspects of the breeding biology of the southern Rockhopper penguin Eudyptes chrysocome chrysocome and new consideration on the intrinsic capacity of the A-egg. Polar Biology 31:925-932.

Putz, K., A. Clausen, N. Huin and J. Croxall.  2003. Re-evaluation of historical Rockhopper penguin population data in the Falkland Islands. Waterbirds 26(2): 169-175.

Putz, K., R. Ingham, J. Smith, and J. Croxall.  2001.  Population trends, breeding success and diet composition of Gentoo Pygoscelis papua, Magellanic Spheniscus magellanicus and RockhopperEudyptes chrysocome penguins in the Falkland Islands.  Polar Biology 24:793–807.

Putz, K., R.J. Ingham, J.G. Smith, and B.H. Luthi. 2002. Winter dispersal of rockhopper penguinsEudyptes chrysocome from the Falkland Islands and its implications for conservation. Marine Ecology Progress Series 240:273-284.

Putz, K., A. Rey, A. Schiavini, A. Clausen, and B. Luthi. 2006. Winter migration of Rockhopper penguins(Eudyptes chrysocome chrysocome) breeding in the Southwest Atlantic: is utilization of different foraging areas reflected in opposing population trends?  Polar Biology 29(9):735–744.

Rey, A.R., and A. Schiavini. 2005. Inter-annual variation in the diet of female southern Rockhopper penguin (Eudyptes chrysocome chrysocome) at Tierra del Fuego. Polar Biology 28:132-141.

Richards, P. 2002. Overview of petroleum geology, oil exploration and associated environmental protection around the Falkland Islands. Aquatic Conservation Marine Freshwater Ecosystems 12:7-14.

Richardson, M.E. 1984. Aspects of ornithology of the Tristan da Cunha group and Gough Island, 1972-1974. Cormorant 12:123-201.

Rounsevell, D.E. and N.P. Brothers. 1984. The status and conservation of seabirds at Macquarie Island. In: Status and Conservation of the World’s Seabirds (eds. J.P. Croxall, P.G.H. Evans and R.W. Schreiber) pp. 587-592. Paston Press, England.

Ryan, P.G. and J. Cooper. 1991. Rockhopper penguins and other marine life threatened by driftnet fisheries at Tristan da Cunha. Oryx 25:76-79.

Ryan, P.G. and C.L. Moloney. 2000. The status of Spectacled Petrels (Procellaria conspicillata) and other sea birds at Inaccessible Island. Marine Ornithology 28:93-100.

Schiavini, A.  2000. Staten Island, Tierra del Fuego: the largest breeding ground for southern Rockhopper Penguins? Waterbirds 23(2):286–291.

Schiavini, A. and A.R. Rey.  2004. Long days, long trips: foraging ecology of female Rockhopper penguins Eudyptes chrysocome chrysocome at Tierra del Fuego. Marine Ecology Progress Series 275:251-262.

Swales, M.K. 1965. The seabirds of Gough Island. Ibis 107:17-42, 215-229.

Taylor, G.A. 2000.  Action Plan for Seabird Conservation in New Zealand. Part A, Threatened Seabirds. Threatened Species Occasional Publication No. 16. New Zealand Department of Conservation, Wellington. Published online at: <>.  [Retrieved May 5, 2008.]  230 pp.

Tristan da Cunha Website. 2009.  Conservation news.  Online at: Retrieved 5 March 2009.

Uhart, M., W. Karesh, R. Cook, N. Huin, K. Lawrence, L. Guzman, H. Pacheco, G. Pizarro, R. Mattsson and T. Morner. 2004. Paralytic shellfish poisoning in Gentoo penguins (Pygoscelis papua) from the Falkland (Malvinas) Islands.  In 2004 proceedings of the AAZV, AAWV, WDA joint conference. pp 481-486.

Verrill, G.E. 1895. On some birds and eggs collected by Mr. George Comer at Gough Island, Kerguelen Island, and the Island of South Georgia, with extracts from his notes, including a meteorological record for about six months at Gough Island. Transactions of the Connecticut Academy of Arts and Sciences 9:430-478.

Watkins, B.P. 1987. Population sizes of King, Rockhopper and Macaroni penguins and Wandering albatrosses at the Prince Edward Islands and Gough Island, 1951-1986. South African Journal of Antarctic Research 17:155-162.

Weimerskirch, H., R. Zotier and P. Jouventin. 1989. The avifauna of the Kerguelen Islands. Emu 89:15-29.

Williams, A. 1981. The clutch size of Macaroni penguins Eudyptes chrysolophus and Rockhopper penguins Eudyptes chrysocome. Emu 81(2): 87.

Williams, T.D. 1995. The Penguins. Oxford University Press.

Williams, A.J. and C. Stone. 1981. Rockhopper penguins Eudyptes chrysocome at Tristan da Cunha. Cormorant 9(2):59-66.

Williams, A.J. and M.J. Imber. 1980. Rockhopper Penguins Eudyptes chrysocome at Gough Island. Bulletin of the British Orinthologists Club 100:208-212.

Woehler, E.J. 1993. The Distribution and Abundance of Antarctic and sub-Antarctic Penguins. Scientific Committee on Antarctic Research; Cambridge. 76 pp.

Woehler, E.J. and J.P. Croxall. 1997.  The status and trends of Antarctic and sub-Antarctic seabirds. Marine Ornithology 25:43-66.